Edited Books

Hill, G. E. and K. J. McGraw. 2006b.  Bird Coloration. Volume II. Function and Evolution. Harvard University Press, Cambridge, MA. Catalog advertisement, Table of Contents, Purchase now, Read an excerpt, Wingspan (review), Times Literary Supplement (review), Science (browsing), Trends in Ecology and Evolution (review), Nature (review), Condor (review), Quarterly Review of Biology (review), ISBE newsletter (review)

Hill, G. E. and K. J. McGraw. 2006a. Bird Coloration. Volume I. Mechanisms and Measurements. Harvard University Press, Cambridge, MA. Catalog advertisement, Table of Contents, Purchase now, ASU Insight feature (on last page), Condor (review), Auk (review)

McGraw Journal Publications and Book Chapters (Google Scholar statistics)

186. Rowe, M., K. Pierson, and K. J. McGraw. 2015. Exploratory behavior is associated with plasma carotenoid accumulation in two congeneric species of waterfowl. Behav. Proc. (in press).

185. Ge, Z., J. D. Johnson, P. A. Cobine, K. J. McGraw, R. Garcia, and G. E. Hill. 2015. High concentrations of ketocarotenoids in the hepatic mitochondria of a molting red songbird. Physiol. Biochem. Zool. (in press).

184. Giraudeau, M., A. Chavez, M. B. Toomey, and K. J. McGraw. 2015. Effects of carotenoid supplementation and oxidative challenges on physiological parameters and carotenoid-based coloration in an urbanization context. Behav. Ecol. Sociobiol. (in press).

183. Fasanello, V. J., E. D. Carlton, M. Pott, N. M. Marchetto, E. Vaughn, K. J. McGraw, R. A. Mauck, and M. F. Haussmann. 2015. Monomorphic ornamentation related to oxidative damage and assortative mating in the black guillemot (Cepphus grylle). Waterbirds 38:106-110.

182. Giraudeau, M., P. M. Nolan, C. Black, S. Earl, M. Hasegawa, and K. J. McGraw. 2015. Song characteristics track bill morphology along a gradient of urbanization in house finches. Frontiers in Zool. (in press).

181. Steffen, J. E., K. M. Learn, J. S. Drumheller, S. M. Boback, and K. J. McGraw. 2015. Carotenoid composition of colorful body stripes and patches in the Painted Turtle (Chrysemys picta) and Red-Eared Slider (Trachemys scripta). Chelonian Cons. Biol. (in press).

180. Pearlstein, E., M. Hughs, J. Mazurek, K. J. McGraw, C. Pesme, and M. Garcia-Garibay. 2014. Correlations between photochemical damage and UV fluorescence of feathers. ICOM-CC Ethnographic Collections (in press).

179. Thomas, D. B., K. J. McGraw, M. W. Butler, M. Carrano, O. Madden, and H. F. James. 2014. Ancient origins and multiple appearances of carotenoid-pigmented feathers in birds. Proc. R. Soc. Lond. B (in press).

178. Newbrey, J., C. Paszkowski, K. J. McGraw, and S. Ogle. 2014. Laying-sequence variation in yolk carotenoids and egg characteristics in the red-winged blackbird, Agelaius phoeniceus. J. Avian Biol. (in press).

177. Ismar, S., C. Daniel, B. Igic, P. Cassey, K. McGraw, D. Dearborn, K. Wakamatsu, A. Odeen, O. Hastad, B. Stephenson, C. Millar, G. Ballard, and M. Hauber. 2014. Sexual plumage dichromatism in a size-monomorphic seabird. Wilson J. Ornithol. (in press).

176. Taylor, L. A., D. Clark, and K. J. McGraw. 2014. Natural variation in condition-dependent display coloration does not predict male courtship success in a jumping spider. Anim. Behav. (in press).

175. Giraudeau, M., and K. J. McGraw. 2014. Physiological correlates of urbanization in a desert songbird. Integr. Comp. Biol. (in press).

174. McCartney, K. L., R. A. Ligon, M. W. Butler, D. F. DeNardo, and K. J. McGraw. 2014. The effect of carotenoid supplementation on immune system development in juvenile male veiled chameleons (Chamaeleo calyptratus). Frontiers in Zoology (in press).

173. Taylor, L. A., D. Clark, and K. J. McGraw. 2014. From spiderling to senescence: ontogeny of color in the jumping spider Habronattus pyrrithrix. J. Arachnol. (in press).

172. Hasegawa, M., R. A. Ligon, M. Giraudeau, M. Watanabe, and K. J. McGraw. 2014. Urban and colorful male house finches are less aggressive. Behav. Ecol. (in press).

171. Friedman, N. R., K. J. McGraw, and K. E. Omland. 2014. History and mechanisms of carotenoid plumage evolution in the New World orioles (Icterus). Comp. Biochem. Physiol. B (in press).

170. Steffen, J. E., K. M. Learn, J. S. Drumheller, S. M. Boback, and K. J. McGraw. 2014. Carotenoid composition of colorful body stripes and patches in the painted turtle (Chrysemys picta) and red-eared slider (Trachemys scripta). J. Herpetol. (in press).

169. Giraudeau, M., M. Mousel, S. Earl, and K. J. McGraw. 2014. Parasites in the city: degree of urbanization predicts poxvirus and coccidian infections in house finches (Haemorhous mexicanus). PLOS One (in press).

168. Thomas, D. B., K. J. McGraw, H. F. James, and O. Madden. 2014. Non-destructive descriptions of carotenoids in feathers using Raman spectroscopy. Analyt. Methods (in press). Link

167. Ligon, R. A. and K. J. McGraw. 2014. Chameleons communicate with complex colour change during contests: different body regions convey different information. Biol. Letters (in press).

166. Peluc, S. I., W. L. Reed, P. Gibbs, and K. J. McGraw. 2014. Maternal dietary carotenoids mitigate detrimental effects of maternal GnRH on offspring immune function in Japanese quail (Coturnix japonica). J. Avian Biol. (in press).

165. Brown, A. C., H. M. Leonard, K. J. McGraw, and E. Clotfelter. 2014.  Maternal effects of carotenoid supplementation in an ornamented cichlid fish (Amantitlania siquia). Funct. Ecol. (in press).

164. McGraw, K. J., M. Giraudeau, G. E. Hill, and M. B. Toomey. 2013. Ketocarotenoid circulation, but not retinal carotenoid accumulation, is linked to eye disease status in a wild songbird. Arch. Biochem. Biophys. 539:156-162.

163. Friedman, N., K. J. McGraw, and K. E. Omland. 2014. Evolution of carotenoid pigmentation in caciques and meadowlarks (Icteridae): repeated gains of red plumage coloration by carotenoid C-4 oxygenation. Evolution (in press).

162. Pickett, S. R. A., S. B. Weber, K. J. McGraw, K. J. Norris, and M. R. Evans. 2013. Environmental and parental influences on offspring health and growth in great tits (Parus major). PLoS One 8:e69786.

161. Butler, M. W., B. Karanfilian, M. Homsher, and K. J. McGraw. 2013. Carotenoid supplementation during adulthood, but not development, decreases testis size in mallards. Comp. Biochem. Physiol. A 166:465-469.

160. Giraudeau, M., K. Sweazea, M. W. Butler, and K. J. McGraw. 2013. Effects of carotenoid and vitamin E supplementation on oxidative stress and plumage coloration in house finches (Haemorhous mexicanus). Comp. Biochem. Physiol. A 166:406-413.

159. Hill, G. E., X. Fu, S. Balenger, K. J. McGraw, M. Giraudeau, and W. R. Hood. 2013. Changes in concentrations of circulating heat shock proteins in House Finches in response to different environmental stressors. J. Field Ornithol. 84:416-424.

158. Alan, R. R., S. R. McWilliams, and K. J. McGraw. 2013. The importance of antioxidants for avian fruit selection during autumn migration. Wilson J. Ornithology 125:513-525.

157. Lendvai, A. Z., M. Giraudeau, J. Nemeth, V. Bako, and K. J. McGraw. 2013. Carotenoid-based plumage coloration reflects feather corticosterone levels in male house finches (Haemorhous mexicanus). Behav. Ecol. Sociobiol. 67:1817-1824.

156. Butler, M. W. and K. J. McGraw. 2013. Eggshell coloration reflects both yolk characteristics and dietary carotenoid history of female mallards. Funct. Ecol. 27:1176-1185.

155. Brown, A. C., K. J. McGraw, and E. Clotfelter. 2013.  Dietary carotenoids increase yellow non-pigment coloration of female convict cichlids (Amantitlania nigrofasciata). Physiol. Biochem. Zool. 86:312-322.

154. Butler, M. W. and K. J. McGraw. 2013. Immune function is related to adult carotenoid and bile pigment levels, but not dietary carotenoid access during development, in female mallard ducks. J. Exp. Biol. 216:2632-2640.

153. Thomas, D. B., McGoverin, C. M., McGraw, K. J., James, H. F., and Madden, O. 2013. Vibrational spectroscopic analyses of unique yellow feather pigments (spheniscins) in penguins. J. R. Soc. Interface 10:20121065.

152. Butler, M. W., Stahlschmidt, Z. R., Ardia, D. R., Davis, S., Davis, J. R., Guillette, Jr., L. J., Johnson, N., McCormick, S. D., McGraw, K. J., and DeNardo, D. F. 2013. Thermal sensitivity of immune function: evidence against a generalist-specialist tradeoff among endothermic and ectothermic vertebrates. Am. Nat. 181:761-774.

151. Taylor, L. A. and K. J. McGraw. 2013. Male ornamental coloration improves courtship success in a jumping spider, but only in the sun. Behav. Ecol. 24:955-967.

150. Cohen, A. A., Bowman, R., Boughton, R. K., Bridge, E., Heiss, R. S., Schoech, S. J., and K. J. McGraw. 2013. Circulating carotenoid levels are negatively associated with previous reproductive success in Florida Scrub-Jays (Aphelocoma coerulescens). Can. J. Zool. 91:64-70.

149. Rowe, M., Tourville, E. A., and K. J. McGraw. 2012. Carotenoids in bird testes: links to body carotenoid supplies, plumage coloration, body mass and testes mass in house finches (Carpodacus mexicanus). Comp. Biochem. Physiol. B 163:285-291. Link

148. Butler, M. W. and K. J. McGraw. 2012. Differential effects of early- and late-life access to carotenoids on adult immune function and ornamentation in mallard ducks (Anas platyrhynchos). PLoS One 7:e38043.

147. Meadows, M. G., Roudybush, T., and K. J. McGraw. 2012. Dietary protein affects iridescent coloration in Anna’s hummingbirds (Calypte anna). J. Exp. Biol. 215:2742-2750.

146. Behbahaninia, H., Butler, M. W., Toomey, M. B., and K. J. McGraw. 2012. Food color preferences against a dark, textured background vary in relation to sex and age in house finches (Carpodacus mexicanus). Behaviour 149:51-65.

145. Toomey, M. B. and K. J. McGraw. 2012. Mate choice for a male carotenoid-based ornament is linked to female dietary carotenoid intake and accumulation. BMC Evol. Biol. 12:3. PDF

144. Peluc, S. I., Reed, W. L, McGraw, K. J., and Gibbs, P. 2012. Carotenoid supplementation and GnRH challenges influence female endocrine physiology, immune function and egg-yolk characteristics in Japanese quail (Coturnix japonica). J. Comp. Physiol. B. 182:687-702.

143. Giraudeau, M., Toomey, M. B., and McGraw, K. J. 2012. Can house finches (Carpodacus mexicanus) use non-visual cues to discriminate the carotenoid content of foods? J. Ornithol. 153:1017-1023.

142. Butler, M. W and K. J. McGraw. 2012. Developmental immune history affects adult immune function but not carotenoid-based ornamentation in mallard ducks. Funct. Ecol. 26:406-415.

141. Toomey, M. B. and K. J. McGraw. 2011. The effects of dietary carotenoid supplementation and retinal carotenoid accumulation on vision-mediated foraging in the house finch. PLoS One 6:e21653. Article

140. Butler, M.W., Toomey, M.B., McGraw, K.J., and Rowe, M. 2011. Ontogenetic immune challenges shape adult personality in mallard ducks. Proc. R. Soc. B 279: 326-333. PDF, ScienceNow piece, Discover magazine piece, Science News, SOLS News

139. Chui, C. K. S., McGraw, K. J., and Doucet, S. M. 2011. Carotenoid-based plumage coloration in golden-crowned kinglets (Regulus satrapa); pigment characterization and relationships with migratory timing and condition. J. Avian Biol. 42:309-322.

138. Walsh, N., Dale, J., McGraw, K.J., Pointer, M. A., and Mundy, N. I. 2012. Candidate genes for carotenoid colouration in vertebrates and their expression profiles in the carotenoid-containing plumage and bill of a wild bird. Proc. R. Soc. B 279:58-66.

137. Rowe, M., Giraudeau, M., Czirjak, G., and McGraw, K. J. 2011. Sexual ornamentation reflects antibacterial activity of ejaculates in mallards. Biology Letters 7:740-742.

136. Lifjeld, J. T., O. Kleven, F. Jacobsen, K. J. McGraw, R. J. Safran, and R. Robertson. 2011. Age before beauty? Relationships between fertilication success and age-dependent ornaments in barn swallows. Behav. Ecol. Sociobiol. 65:1687-1697.

135. Dugas, M. B. and K. J. McGraw. 2011. Nestling mouth coloration: proximate mechanisms, correlates, and constraints in house sparrows (Passer domesticus). Condor 113:691-700.

134. McGraw, K. J., K. Lee, and A. Lewin. 2011. The effect of capture-and-handling stress on carotenoid-based beak coloration in zebra finches. J. Comp. Physiol. A 197:683-691.

133. Navarro, C., T. Perez-Contreras, J. M. Avies, K. J. McGraw, and J. J. Soler. 2011. Blue-green eggshell coloration reflects yolk antioxidant content in spotless starlings (Stumus unicolor). J. Avian Biol. 42:538-543.

132. Meadows, M. G., N. I. Morehouse, R. L. Rutowski, J. M. Douglas, and K. J. McGraw. 2011. Quantifying iridescent coloration in animals: a method for improving repeatability. Behav. Ecol. Sociobiol. 65:1317-1327.

131. Smith, C. L., M. B. Toomey, B. R. Walker, E. J. Braun, B. O. Wolf, K. J. McGraw, and K. L. Sweazea. 2011. High levels of circulating antioxidants protect from the development of oxidative stress in the vasculature of mourning doves (Zenaida macroura). Zoology 114:171-176.

130. Butler, M. W. and K. J. McGraw. 2011. Past or present? Effect of developmental and adult conditions on adult health and coloration in mallards. J. Comp. Physiol. B 181:551-563.

129. Heiss, R. S., A. A. Cohen, R. Bowman, R. K. Boughton, E. Bridge, K. J. McGraw, and S. J. Schoech. 2011. Circulating antioxidant concentrations are negatively correlated with timing of breeding in Florida Scrub-Jays (Aphelocoma coerulescens). J. Exp. Zool. A 315A:101-110.

128. Weiss, S. L., E. A. Kennedy, R. J. Safran, and K. J. McGraw. 2011. Pterin-based ornamental coloration predicts yolk antioxidant levels in female striped plateau lizards (Sceloporus virgatus). J. Anim. Ecol. 80:519-527.

127. Butler, M. W., M. B. Toomey, and K. J. McGraw. 2011. How many color metrics do we need? Evaluating how different color-scoring procedures explain carotenoid pigment content in avian bare-part and plumage ornaments. Behav. Ecol. Sociobiol. 65:401-413.

126. Taylor, L. A., D. L. Clark, and K. J. McGraw. 2011. Condition-dependence of male display coloration in a jumping spider (Habronattus pyrrithrix). Behav. Ecol. Sociobiol. 65:1133-1146.

125. McGraw, K. J., O. L. Crino, and P. M. Nolan. 2011. Carotenoids boost immunity during molt in a songbird with sexually selected carotenoid-based plumage coloration. Biol. J. Linn. Soc. 102:560-572.

124. Burtt, Jr., E. H., M. R. Schroeder, L. A. Smith, J. E. Sroka, and K. J. McGraw. 2011. Colourful parrot feathers resist bacterial degradation. Biol. Letters 7:214-216. PDF

123. Giraudeau, M., C. Duval, G. A. Czirjak, V. Bretagnolle, C. Eraud, K. J. McGraw, and P. Heeb. 2011. Maternal investment of female mallards is influenced by male carotenoid-based coloration. Proc. R. Soc. B 278:781-788. PDF

122. Giraudeau, M., G. A. Czirjak, C. Duval, V. Bretagnolle, C. Eraud, K. J. McGraw, and P. Heeb. 2011. Effect of restricted pree-gland access on maternal self maintenance and reproductive investment in mallards. PLoS One 5:e13555. PDF

121. McGraw, K. J., A. Cohen, D. Costantini, and P. Horak. 2010. The ecological significance of antioxidants and oxidative stress: a marriage of functional and mechanistic approaches. Funct. Ecol. (invited editorial) 24: 947-949. PDF

120. Butler, M. W. and K. J. McGraw. 2010. Relationships between dietary carotenoids, body tissue carotenoids, parasite burden, and health state in wild mallard (Anas platyrhynchos) ducklings. Arch. Biochem. Biophys. 504:154-160. PDF

119. Costantini, D., M. Rowe, M. W. Butler, and K. J. McGraw. 2010. From molecules to living systems: historical and contemporary issues in oxidative stress and antioxidant ecology. Funct. Ecol. (invited review) 24:950-959. PDF

118. Toomey, M. B. and K. J. McGraw. 2010. The effects of dietary carotenoid intake on carotenoid accumulation in the retina of a wild bird, the house finch (Carpodacus mexicanus). Arch. Biochem. Biophys. 504:161-168. PDF

117. Hipfner, J. M., J. Dale, and K. J. McGraw. 2010. Yolk carotenoids and stable isotopes reveal links among environment, foraging behaviour and seabird breeding success. Oecologia 163:351-360.

116. Safran, R. J., K. J. McGraw, K. M. Pilz, and S. M. Correa. 2010. Egg-yolk androgen and carotenoid deposition as a function of maternal social environment in barn swallows Hirundo rustica. J. Avian Biol. 41:470-478.

115. Toomey, M. B., M. W. Butler, and K. J. McGraw. 2010. Immune-system activation depletes retinal carotenoids in house finches. J. Exp. Biol. 213:1709-1716.

114. Nolan, P. M., F. S. Dobson, T. J. Karels, K. J. McGraw, and P. Jouventin. 2010. Mutual mate choice for colorful traits in king penguins. Ethology 116:635-644.

113. Toomey, M. B., M. W. Butler, M. G. Meadows, L. A. Taylor, H. Fokidis, and K. J. McGraw. 2010. A novel method for quantifying the glossiness of animals. Behav. Ecol. Sociobiol. 64:1047-1055. Uncorrected page proofs

112. Navarro Puig, C., T. Perez-Contreras, J. Aviles, J. Soler, and K. J. McGraw. 2010. Beak colour reflects circulating carotenoid and vitamin A levels in spotless starlings (Sturnus unicolor). Behav. Ecol. Sociobiol. .

111. McGraw, K. J. 2010. Avian antioxidants and oxidative stress: insights from food, physiology, and feathers. In: Oxidative stress (Mandelker, L. and Vajdovich, P., eds), Springer, NY.

110.  Hipfner, J. M., K. A. Hobson, J. Dale, and K. J. McGraw. 2010. Stable isotopes link diet to avian yolk carotenoid allocation: a comparative study of five auk species (Charadriiformes: Alcidae). Physiol. Biochem. Zool. (in press).

109. Safran, R. J., K. J. McGraw, M. R. Wilkins, J. K. Hubbard, and J. Marling. 2010. Positive carotenoid balance correlates with greater reproductive performance in a wild bird. PLoS One 5(2): e9420. doi:10.1371/journal.pone.0009420 PDF, press coverage

108. Lin, S. M., K. Nieves-Puigdoller, A. C. Brown, K. J. McGraw, and E. D. Clotfelter. 2010. Testing the carotenoid trade-off hypothesis in the polychromatic Midas cichlid, Amphilophus citrinellus. Physiol. Biochem. Zool. 83:333-342. PDF

107. McGraw, K. J. and M. B. Toomey. 2010. Carotenoid accumulation in the tissues of zebra finches: predictors of integumentary pigmentation and implications for carotenoid allocation strategies. Physiol. Biochem. Zool. 83:97-109. PDF and appendix

106. Cohen, A. A., R. A. Mauck, N. T. Wheelwright, C. E. Huntington, and K. J. McGraw. 2009. Complexity in relationships between antioxidants and individual life-history parameters in a songbird and a seabird. Oikos 118:1854-1861. PDF

105. McGraw, K. J. and J. D. Blount. 2009. Control and function of carotenoid coloration in birds: a review of case studies. In Carotenoids: Physical, Chemical, and Biological Functions and Properties (J. T. Landrum, ed.). CRC Press, Boca Raton, FL (in press; published 21 December 2009).

104. Butler, M. W. and K. J. McGraw. 2009. Indoor housing during development affects moult, carotenoid circulation, and beak colouration of mallard ducks (Anas platyrhynchos). Avian Biol. Res. 2:203-211. PDF

103. Dierenfeld, E.S., K. J. McGraw, K. Fritsche, J.T. Briggler, and J. Ettling. 2009. Nutrient composition of whole crayfish (Orconectes and Procambarus species) consumed by hellbender (Cryptobranchus alleganiensis). Herp. Rev. 40:324-330. PDF

102. Cohen, A. A., W. D. Robinson, and K. J. McGraw. 2009. Serum antioxidant levels in wild birds vary in relation to diet, season, life history strategy, and species. Oecologia 161:673-683. PDF, Supplementary Material 1, Supplementary Material 2

101. Bascunan, A. L., E. A. Tourville, M. B. Toomey, and K. J. McGraw. 2009. Food color preferences of molting house finches (Carpodacus mexicanus) in relation to sex and plumage coloration. Ethology 115:1066-1073. PDF

100. Steffen, J. and K. J. McGraw. 2009. How dewlap color reflects its carotenoid and pterin content in brown anoles. Comp. Biochem. Physiol. B 154:334-340. PDF

99. McGraw, K. J. 2009. Identifying anatomical sites of carotenoid metabolism in birds. Naturwissenschaften 96:987-988. PDF, Original article critiqued, Authors’ response to my critique

98. McGraw, K. J., M. Massaro, T. J. Rivers, and T. Mattern. 2009. Annual, sexual, size-, and condition-related variation in the colour and fluorescent pigment content of yellow crests in Snares penguins (Eudyptes robustus). Emu 109:93-99. PDF, cover image

97. McGraw, K. J. 2009. Canine coloration: dark domesticated dogs and the bright side for wild wolves. Pigment Cell Melanoma Res. 22:255-256. PDF

96. Meadows, M. G., M. W. Butler, N. I. Morehouse, L. A. Taylor, M. B. Toomey, K. J. McGraw, and R. L. Rutowski. 2009. Iridescence: views from many angles. J. R. Soc. Interface 6:S107-S113. PDF

95. Cohen, A. A. and K. J. McGraw. 2009. No simple measures for antioxidant status in birds: complexity in inter- and intraspecific correlations among circulating antioxidant types. Funct. Ecol. 23:310-320. PDF

94. Toomey, M. B. and K. J. McGraw. 2009. Seasonal, sexual, and quality related variation in retinal carotenoid accumulation in the house finch (Carpodacus mexicanus). Funct. Ecol. 23:321-329. PDF

93. McGraw, K. J. and A. L. A. Middleton. 2009. American Goldfinch (Carduelis tristis). The Birds of North America Online (A. Poole, Ed.). Ithaca, NY: Cornell Laboratory of Ornithology.

92. Rowe, M. and K. J. McGraw. 2009. Carotenoids in the seminal fluid of wild birds: interspecific variation in fairy-wrens. Condor 110:694-700. PDF

91. McGraw, K. J. 2009. Visual signaling in animals. In: Encyclopedia of Neuroscience (L. R. Squire, ed.), vol. 10. Academic Press Oxford, pp. 307-311. PDF

90. McGraw, K. J., E. A. Tourville, and M. W. Butler. 2008. A quantitative comparison of the commonly used methods for extracting carotenoids from avian plasma. Behav. Ecol. Sociobiol. 62:1991-2002. PDF

89. Cohen, A. A., K. J. McGraw, P. Wiersma, J. B. Williams, W. D. Robinson, T. R. Robinson, J. D. Brawn, and R. E. Ricklefs. 2008. Interspecific associations between circulating antioxidant levels and life-history variation in birds. Am. Nat. 172:178-193. PDF

88. Piault, R., J. Gasparini, P. Bize, M. Paulet, K. J. McGraw, and A. Roulin. 2008. Experimental support for the make-up hypothesis in nestling tawny owls (Strix aluco). Behav. Ecol. 19:703-709. PDF

87. Safran, R. J., J. Adelman, K. J. McGraw, and M. Hau. 2008. Sexual signal elaboration affects physiological state in a social vertebrate. Current Biology 18:R461-R462. PDF, over 300 newspaper and online articles written on this paper, plus Good Morning America, CNN, and several local radio interviews (e.g. San Francisco, Houston, Rockford, IL, New York). TREE “Research Focus” article highlighting this paper

86. Deviche, P., K. J. McGraw, and J. Underwood. 2008.  Age-, sex- and season-specific accumulation of plasma carotenoid pigments in white-winged crossbills. J. Avian Biol. 39:283-292. PDF

85. Blount, J. D. and K. J. McGraw. 2008. Signal functions of carotenoid colouration. In: Carotenoids. Volume 4: Natural Functions (G. Britton, S. Liaaen-Jensen, and H. Pfander, eds.). Birkhauser Verlag, basel, pp. 213-236. Uncorrected galley proofs

84. McGraw, K. J. 2008. An update on the honesty of melanin-based color signals in birds. Pigment Cell Melanoma Res. 21:133-138. PDF

83. Schaefer, H. M., K. J. McGraw, and C. Catoni. 2008. Birds use fruit colour as an honest signal of dietary antioxidant rewards. Funct. Ecol. 22:303-310. PDF

82. Juola, F. A., K. J. McGraw, and D. C. Dearborn. 2008. Carotenoids and throat pouch coloration in the great frigatebird (Fregata minor). Comp. Biochem. Physiol. B. 149:370-377. PDF

81. Safran, R. J., Pilz, K. M., McGraw, K. J., Correa, S. M., and Schwabl, H. 2008. Are yolk androgens and carotenoids in barn swallow eggs related to parental quality? Behav. Ecol. Sociobiol. 62:427-438. PDF

80. Jouventin, P., K. J. McGraw, M. Morel, and A. Celerier. 2008. Dietary carotenoid supplementation affects orange beak but not foot coloration in gentoo penguins. Waterbirds 30:573-578. PDF

79. Clotfelter, E., Ardia, D., and McGraw, K. J. 2007. Red fish, blue fish: trade-offs between pigmentation and immunity in Betta splendens. Behav. Ecol. 18:1139-1145. PDF

78. Toomey, M. B. and K. J. McGraw. 2007. Modified saponification and HPLC methods for analyzing carotenoids from the retina of Japanese quail (Coturnix japonica): implications for its use as a nonprimate model species. Invest. Ophthalmol. Vis. Sci. 48:3976-3982. PDF

77. Taylor, L. A. and K. J. McGraw. 2007. Animal coloration: sexy spider scales. Curr. Biol. 17:R592-R593. PDF

76. McGraw, K. J., M. B. Toomey, P. M. Nolan, N. I. Morehouse, M. Massaro, and P. Jouventin. 2007. A description of unique fluorescent yellow pigments in penguin feathers. Pigment Cell Res. 20:301-304. PDF, Supplementary Material

75. Bezzerides, A. L., K. J. McGraw, R. S. Parker, and J. Husseini. 2007. Elytra color as a signal of chemical defense in the Asian ladybird beetle (Harmonia axyridis). Behav. Ecol. Sociobiol. 61:1401-1408. PDF

74. McGraw, K. J. and D. R. Ardia. 2007. Do carotenoids buffer testosterone-mediated immunosuppression?: an experimental test in a colorful songbird. Biol. Letters 3:375-378. PDF, Electronic Supplementary Material, Biology Letters Highlighted Paper, ASU Headline, ASU News, School of Life Sciences News, Science Daily, Innovations Report, Biology News Net,, Eureka Alert,, United Press International, ImediNews, Earth Times, Monsters and Critics, New Kerala, Post Chronicle,, Associated Content, Medical News Today, MediLexicon, Nutrition Horizon, ASU Research Magazine, Softpedia

73. Hofmann, C. M., K. J. McGraw, T. W. Cronin, and K. E. Omland. 2007. Melanin coloration in New World orioles I: carotenoid masking and pigment dichromatism in the orchard oriole complex. J. Avian Biol. 38:163-171. PDF

72. McGraw, K. J., W. Medina-Jerez, and H. Adams. 2007. Carotenoid-based plumage coloration and aggression during molt in male house finches. Behaviour 144:165-178. PDF

71. Steffen, J. and K. J. McGraw. 2007. Patterns of carotenoid and pterin pigmentation in the colorful dewlaps of two anole species. Comp. Biochem. Physiol. B. 146:42-46. PDF

70. McGraw, K. J. 2007. Dietary mineral content influences melanin-based ornamental coloration. Behav. Ecol. 18:137-142. PDF

69. McGraw, K. J. 2006. The mechanics of uncommon colors in birds: pterins, porphyrins, and psittacofulvins. In: Bird Coloration. I. Mechanisms and Measurements (G. E. Hill and K. J. McGraw, eds.). Harvard University Press, Cambridge, MA, pp. 354-398.      

68. McGraw, K. J. 2006. The mechanics of carotenoid coloration in birds. In: Bird Coloration. I. Mechanisms and Measurements (G. E. Hill and K. J. McGraw, eds.). Harvard University Press, Cambridge, MA, pp. 177-242.

67. McGraw, K. J. 2006. The mechanics of melanin coloration in birds. In: Bird Coloration. I. Mechanisms and Measurements (G. E. Hill and K. J. McGraw, eds.). Harvard University Press, Cambridge, MA, pp. 243-294.

66. Shawkey, M. D., G. E. Hill, K. J. McGraw, and W. R. Hood. 2006. An experimental test of the contributions and condition-dependence of microstructure and carotenoids in yellow plumage colouration. Proc. R. Soc. Lond. B. 273:2985-2991. PDF

65. McGraw, K. J., O. L. Crino, W. Medina-Jerez, and P. M. Nolan. 2006. Effect of dietary carotenoid supplementation on food intake and immune function in a songbird with no carotenoid coloration. Ethology 112: 1209-1216. PDF

64. McGraw, K. J. and K. C. Klasing. 2006. Carotenoids, immunity, and integumentary coloration in red junglefowl (Gallus gallus). Auk 123:1161-1171. PDF

63. McGraw, K. J. 2006. Carotenoids mediate a trade-off between egg quantity and quality in Japanese quail. Ethol. Ecol. Evol. 18:247-256. PDF

62. McGraw, K. J. 2006. Sex-steroid dependence of carotenoid-based coloration in female zebra finches. Physiol. Behav. 88:347-352. PDF

61. McGraw, K. J., P. M. Nolan, and O. L. Crino. 2006. Carotenoid accumulation strategies for becoming a colorful house finch: analyses of plasma and liver pigments in wild molting birds. Funct. Ecol. 20:678-688. PDF, Cover Image

60. McGraw, K. J., S. M. Correa, and E. Adkins-Regan. 2006. Testosterone upregulates lipoprotein status to control sexual attractiveness in a colorful songbird. Behav. Ecol. Sociobiol. 60:117-122. PDF

59. McGraw, K. J. and L. Hardy. 2006. Astaxanthin is responsible for the pink plumage flush of Franklins and Ring-billed gulls. J. Field Ornithol. 77:29-33. PDF

58. McGraw, K. J. and R. S. Parker. 2006. A novel lipoprotein-mediated mechanism controlling sexual attractiveness in a colorful songbird. Physiol. Behav. 87:103-108. PDF

57. Deviche, P., K. McGraw, and E. Greiner. 2005. Interspecific differences in hematozoan infection in Sonoran Desert Aimophila sparrows. J. Wild. Dis. 41:532-541. PDF

56. McGraw, K. J., E. Adkins-Regan, and R. S. Parker. 2005. Maternally derived carotenoid pigments affect offspring survival, sex ratio, and sexual attractiveness in a colorful songbird. Naturwiss. 92:375-380. PDF

55. McGraw, K. J., R. J. Safran, and K. Wakamatsu. 2005. How feather colour reflects its melanin content. Funct. Ecol. 19:816-821. PDF

54. Safran, R. J., C. R. Neuman, K. J. McGraw, and I. J. Lovette. 2005. Dynamic paternity allocation as a function of male plumage color in barn swallows. Science 309:2210-2212. PDF, Cover image, Cornell University, Arizona State University, ASU Insight, ASU homepage, BBC, Cornell Daily Sun, Cornell Lab of Ornithology, American Association for the Advancement of Science, ABC News, ABC News Online, AOL News, Belleville News Democrat, Biology News Net, Cherry Bomb, CLAS Alumni Magazine, EurekAlert, East Valley Tribune (AZ), East Valley Tribune homepage, Melbourne Herald Sun, Innovations Report, Independent Online, Live Science, MSNBC, National Geographic, National Wildlife Federation Magazine, New Haven Herald, New Kerala, New Scientist, New York Times, Newswise, Noorderlicht, NPR, OptusNet, Pet Talk Radio, Planet Ark, Red Nova, Reuters, Salem Statesman Journal, Science Daily, School of Life Sciences (ASU), Science News, Telegraph, The Age, The Oregonian, Times Online, Unison, Virtual Life Sciences Library, Web India, World News Australia, Yahoo News, AAAS radio clip, CLAS News (ASU), 10,000 Birds

53. McGraw, K. J. 2005. Interspecific variation in dietary carotenoid assimilation in birds: links to phylogeny and color ornamentation. Comp. Biochem. Physiol. B 142:245-250. PDF

52. McGraw, K. J. and M. C. Nogare. 2005. Distribution of unique red feather pigments in parrots. Biology Letters 1:38-43. PDF, Arizona State University homepage 1, Arizona State University homepage 2, Arizona State University press release 1, Arizona State University press release 2, Arizona State University video interview, ASU Insight Newspaper, The Arizona Republic, Innovations Report, The Biochemist, Windy City Parrot, Wissenschaft, EurekaAlert (AAAS), ChemLin, ParrotSuperCenter, Yahoo Korea, Noorderlicht, ScienceDaily, LiveScience,, The Oregonian, Farbimpulse, Futura Sciences, El Correo Digital, Vigyan Prasar, Precious Fids, Girl Scientist, Science News, Aviary & Cage Bird Society of South Florida, ASU Insight (Summer Highlights), ASU Alumni Magazine (Spring 2005), eNewsSource, BBC Wildlife, BirdTalk Magazine, Grade Winner, Planned Parrothood, Chemical and Engineering News, Birdwatchers Digest

51. McGraw, K. J. 2005. The antioxidant function of many animal pigments: are there consistent health benefits of sexually selected colorants? Anim. Behav. 69:757-764. PDF, Top 10 article in Animal Behaviour

50. McGraw, K. J. and D. R. Ardia. 2005. Sex differences in carotenoid status and immune performance in zebra finches. Evol. Ecol. Res. 7:251-262. PDF

49. McGraw, K. J., G. E. Hill, and R. S. Parker. 2005. The physiological costs of being colourful: nutritional control of carotenoid utilization in the American goldfinch (Carduelis tristis). Anim. Behav. 69:653-660. PDF

48. McGraw, K. J., J. Hudon, G. E. Hill, and R. S. Parker. 2005. A simple and inexpensive chemical test for behavioral ecologists to determine the presence of carotenoid pigments in animal tissues. Behav. Ecol. Sociobiol. 57:391-397. PDF

47. McGraw, K. J. 2004. Not all red, orange, and yellow animal colors are carotenoid-based: the need to couple biochemical and behavioral studies of color signals. Proc. Ind. Natl. Sci. Acad. B 70:593-598. PDF

46. McGraw, K. J., K. Wakamatsu, A. B. Clark, and K. Yasukawa. 2004. Red-winged blackbirds Agelaius phoeniceus use carotenoid and melanin pigments to color their epaulets. J. Avian Biol. 35:543-550. PDF

45. McGraw, K. J. 2004. Colorful songbirds metabolize carotenoids at the integument. J. Avian Biol. 35:471-476. PDF

44. McGraw, K. J. and A. J. Gregory. 2004. Carotenoid pigments in male American goldfinches: what is the optimal biochemical strategy for becoming colourful? Biol. J. Linn. Soc. 83:273-280. PDF

43. McGraw, K. J. and J. G. Schuetz. 2004. The evolution of carotenoid coloration in estrildid finches: a biochemical analysis. Comp. Biochem. Physiol. B 139:45-51. PDF

42. McGraw, K. J. 2004. Winter plumage coloration in male American goldfinches: do reduced ornaments serve signaling functions in the non-breeding season? Ethology 110:707-715. PDF

41. McGraw, K. J., G. E. Hill, K. J. Navara, and R. S. Parker. 2004. Differential accumulation and pigmenting ability of dietary carotenoids in colorful finches. Physiol. Biochem. Zool. 77:484-491. PDF

40. McGraw, K. J. and K. Wakamatsu. 2004. Melanin basis of ornamental feather colors in male zebra finches. Condor 106:686-690. PDF

39. Horak, P., L. Saks, U. Karu, I. Ots, P. F. Surai, and K. J. McGraw. 2004. How coccidian parasites affect health and appearance of greenfinches. J. Anim. Ecol. 73:935-947. PDF

38. McGraw, K. J., R. J. Safran, M. R. Evans, and K. Wakamatsu. 2004. European barn swallows use melanin pigments to color their feathers brown. Behav. Ecol. 15:889-891. PDF, Confirmatory reply by Stradi

37. McGraw, K. J. and G. E. Hill. 2004. Plumage color as a dynamic trait: carotenoid pigmentation of male house finches (Carpodacus mexicanus) fades during the breeding season. Can. J. Zool. 82:734-738. PDF

36. McGraw, K. J. and M. C. Nogare. 2004. Carotenoid pigments and the selectivity of psittacofulvin-based coloration systems in parrots. Comp. Biochem. Physiol. B 138:229-233. PDF

35. Safran, R. J. and K. J. McGraw. 2004. Plumage coloration, not length or symmetry of tail-streamers, is a sexually selected trait in North American barn swallows. Behav. Ecol. 15:455-461. PDF, Nature Australia

34. McGraw, K. J., K. Wakamatsu, S. Ito, P. M. Nolan, P. Jouventin, F. S. Dobson, R. E. Austic, R. J. Safran, L. M. Siefferman, G. E. Hill, and R. S. Parker. 2004. You cant judge a pigment by its color: carotenoid and melanin content of yellow and brown feathers in swallows, bluebirds, penguins, and domestic chickens. Condor 106:390-395. PDF

33. McGraw, K. J. 2004. Multiple UV reflectance peaks in the iridescent neck feathers of pigeons. Naturwiss. 91:125-129. PDF

32. Mays Jr., H. L., K. J. McGraw, G. Ritchison, S. Cooper, V. Rush, and R. S. Parker. 2004. Sexual dichromatism in the Yellow-breasted Chat (Icteria virens): spectrophotometric analysis and biochemical basis. J. Avian Biol. 35:125-134. PDF

31. McGraw, K. J. and G. E. Hill. 2004. Mate attentiveness, seasonal timing of breeding and long-term pair bonding in the house finch (Carpodacus mexicanus). Behaviour 141:1-13. PDF

30. McGraw, K. J. and D. R. Ardia. 2004. Immunoregulatory activity of different dietary carotenoids in male zebra finches. Chemoecol. 14:25-29. PDF

29. Hill, G. E. and K. J. McGraw. 2004. Correlated changes in male plumage coloration and female mate choice in cardueline finches. Anim. Behav. 67:27-35. PDF

28. McGraw, K. J. and D. R. Ardia. 2003. Carotenoids, immunocompetence, and the information content of sexual colors: an experimental test. Am. Nat. 162:704-712. PDF

27. Saks, L., K. J. McGraw, and P. Horak. 2003. How feather colour reflects its carotenoid content. Funct. Ecol. 17:555-561. PDF

26. McGraw, K. J., M. D. Beebee, G. E. Hill, and R. S. Parker. 2003. Lutein-based plumage coloration in songbirds is a consequence of selective pigment incorporation into feathers. Comp. Biochem. Physiol. B. 135:689-696. PDF

25. McGraw, K. J., G. E. Hill, and R. S. Parker. 2003. Carotenoid pigments in a mutant cardinal: implications for the genetic and enzymatic control mechanisms of carotenoid metabolism in birds. Condor 105:587-592. PDF

24. McGraw, K. J., A. J. Gregory, R. S. Parker, and E. Adkins-Regan. 2003. Diet, plasma carotenoids, and sexual coloration in the zebra finch (Taeniopygia guttata). Auk 120:400-410. PDF

23. McGraw, K. J. 2003. Melanins, metals, and mate quality. Oikos 102:402-406. PDF

22. Hill, G. E and K. J. McGraw. 2003. Melanin, nutrition, and the lions mane. Science (Letter) 299:660. PDF

21. McGraw, K. J., J. Dale, and E. A. Mackillop. 2003. Social environment during molt and the expression of melanin-based plumage pigmentation in male house sparrows  (Passer domesticus). Behav. Ecol. Sociobiol. 53:116-122. PDF

20. McGraw, K. J., E. A. Mackillop, J. Dale, and M. E. Hauber. 2002. Different colors reveal different information: how nutritional stress affects the expression of melanin- and structurally based ornamental coloration. J. Exp. Biol. 205:3747-3755. PDF, JEB 2002 Highlight

19. McGraw, K. J., E. Adkins-Regan, and R. S. Parker. 2002. Anhydrolutein in the zebra finch: a new, metabolically derived carotenoid in birds. Comp. Biochem. Physiol. B. 132:811-818. PDF

18. McGraw, K. J. 2002. Environmental predictors of geographic variation in human mating preferences. Ethology 108:303-317. PDF, Reuters press release, New Scientist article, Straits Times, Ananova, iAfrica, Daily Dispatch, NY Post editorial, Rochester (N.Y.) Democrat & Chronicle, Yahoo News (in Spanish), Expressen (Swedish newspaper), Het Parool (Dutch daily newspaper), Ithaca (N.Y.) Journal, Atlanta Journal-Constitution, Cornell Univ. press release, Australian Broadcasting Company, Hindustan Times (India), Megastar (England), La Opinion (Los Angeles), Washington Post, Hola Hoy (NYC Spanish newspaper), Chicago Sun-Times, Dagbladet (Norwegian newspaper), Indianapolis Star, Terra Lycos, Telegraph News (U.K.), Cornell Chronicle, National Review, Some girl named Allison, San Francisco Chronicle, Indianapolis Star again, Chicago Tribune, Indianapolis Star yet again, Buffalo News, New Orleans Channel, Financial Times magazine (London), EurekAlert (AAAS), Cosmiverse, InteliHealth, El Norte, El Universal, KLIK Magazin, GIGA (Denmark), Newswise, Mindful-Things, SexNews Daily, The KCRA Channel (California), Grinning Idiot, Venezuela Innovadora, Tiscali (Netherlands), Tercera (Chile), Shape Magazine, Allure Magazine, Cornell Communique, Cornell Alumni Magazine, YupiMSN, Avisa Norland (Norway), Aftonbladet (Sweden), 3sat Online (Denmark), Modern Bride magazine, Most Frequently Dowloaded Ethology paper in 2003

17. McGraw, K. J., G. E. Hill, R. Stradi, and R. S. Parker. 2002. The effect of dietary carotenoid access on sexual dichromatism and plumage pigment composition in the American goldfinch. Comp. Biochem. Physiol. B 131:261-269. PDF

16. Badyaev, A. V., Hill, G. E., Beck, M. L., Dervan, A. A., Duckworth, R. A., McGraw, K. J., Nolan, P. M., and Whittingham, L. A.  2002.  Sex-biased hatching order and adaptive population divergence in a passerine bird.  Science 295:316-318. PDF, News of the Week, USA Today, National Geographic News.

15. McGraw, K. J. and G. E. Hill. 2002. Testing reversed sexual dominance from an ontogenetic perspective: juvenile female House Finches Carpodacus mexicanus are dominant to juvenile males. Ibis 144:39-42. PDF

14. McGraw, K. J., P. M. Nolan, A. M. Stoehr, and G. E. Hill. 2001. Intersexual differences in age-specific parental effort in the house finch. Etologia 9:35-41. PDF, Journal cover

13. McGraw, K. J., G. E. Hill, R. Stradi, and R. S. Parker. 2001. The influence of carotenoid acquisition and utilization on the maintenance of species-typical plumage pigmentation in male American goldfinches (Carduelis tristis) and northern cardinalis (Cardinalis cardinalis). Physiol. Biochem. Zool. 74:843-852. PDF, Online version with color photos

12. McGraw, K. J. and G. E. Hill. 2001. Carotenoid access and intraspecific variation in plumage pigmentation in male American goldfinches (Carduelis tristis) and northern cardinals (Cardinalis cardinalis). Funct. Ecol. 15:732-739. PDF, Journal cover

11. McGraw, K. J., A. M. Stoehr, P. M. Nolan, and G. E. Hill. 2001. Plumage redness predicts breeding onset and reproductive success in the house finch: a validation of Darwin's theory. J. Avian Biol. 32:90-94. PDF

10. Stoehr, A. M. and K. J. McGraw. 2001. Ultraviolet reflectance of color patches in male Sceloporus undulatus and Anolis carolinensis. J. Herpetol. 35:168-171. PDF

9. Stoehr, A. M, K. J. McGraw, P. M. Nolan, and G. E. Hill. 2001. Parental care in relation to brood size in the house finch. J. Field Ornithol. 72:412-418. PDF

8. Nolan, P. M., A. M. Stoehr, G. E. Hill and K. J. McGraw. 2001. The number of provisioning visits by house finches predicts the mass of food delivered. Condor 103:851-855. PDF

7. McGraw, K. J. and G. E. Hill. 2000. Plumage brightness and breeding-season dominance in the house finch: a negatively correlated handicap? Condor 102:457-462. PDF

6. McGraw, K. J. and G. E. Hill. 2000. Differential effects of endoparasitism on the expression of carotenoid- and melanin-based ornamental coloration. Proc. R. Soc. Lond. B. 267:1525-1532. PDF, NPR Radio Show, Birdwatcher’s Digest

5. McGraw, K. J. and G. E. Hill. 2000. Carotenoid-based ornamentation and status signaling in the house finch. Behav. Ecol. 11: 520-527. PDF

4. Stoehr, A. M., P. M. Nolan, G. E. Hill, and K. J. McGraw. 2000. Nest mites (Pellonyssus reedi) and the reproductive biology of the house finch (Carpodacus mexicanus). Can. J. Zool. 78:2126-2133. PDF

3. Badyaev, A. V., G. E. Hill, A. M. Stoehr, P. M. Nolan, and K. J. McGraw. 2000. The evolution of sexual dimorphism in the house finch: II. Population divergence in relation to local selection. Evolution 54:2134-2144. PDF

2. McGraw, K. J. and G. E. Hill. 1999. Induced homosexual behaviour in male house finches (Carpodacus mexicanus): the "Prisoner Effect". Ethol. Ecol. Evol. 11:197-201. PDF

1. McGraw, K. J., G. E. Hill and A. J. Keyser. 1999. Ultraviolet reflectance of colored plastic leg bands. J. Field Ornithol. 70:236-243. PDF

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